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A thematic course exploring five fundamental features of all biological systems, including structure and function, information flow, energy and matter, interactions, and evolution. This course examines the integrative biology of coloration, a feature shared by nearly all animals on earth that can be critical to survival and reproduction. Topics discussed in this class include the genetic and cellular underpinnings of coloration, the functions of coloration, how coloration interacts with other traits and processes (e.g., physiology, behavior, and morphology), and the evolutionary reasons why colors may arise, persist, and/or vary across space and time.